The earliest mammal of the Euopean Paloecene: the multituberculate Hainina

Abstract

ABSTRACT—A new species of multituberculate mammal, Hainina pyrenaica n. sp. is described from Fontllonga-3 (Tremp Basin,
Southern Pyrenees, Spain), correlated to the later part of chron C29r just above the K/T boundary. This taxon represents the earliest
European Tertiary mammal recovered so far, and is related to other Hainina species from the European Paleocene. A revision of the
species of Hainina allows recognition of a new species, H. vianeyae n. sp. from the Late Paleocene of Cernay (France). The genus is
included in the family Kogaionidae Ra˜dulescu and Samson, 1996 from the Late Cretaceous of Romania on the basis of unique dental
characters. The Kogaionidae had a peculiar masticatory system with a large, blade-like lower p4, similar to that of advanced Ptilodontoidea,
but occluding against two small upper premolars, interpreted as P4 and P5, instead of a large upper P4. The endemic European
Kogaionidae derive from an Early Cretaceous group with five premolars, and evolved during the Late Cretaceous and Paleocene. The
genus Hainina represents a European multituberculate family that survived the K/T boundary mass extinction event